Jack Klugman, who made an art of gruffness in TV's The Odd Couple and Quincy, ME, has died at the age of 90.

The actor's son Adam says his father died on Monday afternoon in Los Angeles.

In the 1970s sitcom The Odd Couple, Klugman played sloppy sports writer Oscar to co-star Tony Randall's Felix, a fussy photographer.

In Quincy, ME, which aired in the US from 1976 to 1983, Klugman played an idealistic, tough-minded medical examiner.

Klugman lost his voice to throat cancer in the 1980s but trained himself to speak again. He returned to acting in a 1993 Broadway revival of Three Men on a Horse.

Klugman split his time between TV, movies and the New York stage.