A group of chimpanzees has been captured on camera apparently lost in grief for the death of a friend.
The apes at the Sanaga-Yong Chimpanzee Rescue Centre in Cameroon watched in silence from their enclosure as the body of Dorothy was wheeled past for burial.
Dorothy was in her 40s and died of heart failure. She was described as a "prominent figure" within the group.
The photo was taken by centre worker Monica Szczupider.
"We were burying Dorothy," she said.
"We brought her in the wheelbarrow to let the others see.
"I cannot emphasise enough how silent and still they were."
She said chimpanzees are typically not like that.
"They are loud, with short attention spans.
"It was unbelievably emotional."
The photo will appear in November's National Geographic magazine.
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